AFP548

SMTP rejecting incoming mail – no relay access…

Help. This did work, and now doesn't, and I cannot identify what broke it.

(I have changed my domain name to LOCALDOMAIN in the text quoted below. the real domain is a lowercase legitimate real domain.)
the message in the logs is:

Nov 2 11:18:57 Evergreen postfix/smtpd[3712]: connect from hole.macnn.com[216.22.45.54]
Nov 2 11:18:57 Evergreen postfix/smtpd[3712]: NOQUEUE: reject: RCPT from hole.macnn.com[216.22.45.54]: 554 : Relay access denied; from= to= proto=SMTP helo=
Nov 2 11:18:58 Evergreen postfix/smtpd[3712]: disconnect from hole.macnn.com[216.22.45.54]

The postconf -n is:

alias_maps = hash:/etc/aliases
command_directory = /usr/sbin
config_directory = /etc/postfix
content_filter = smtp-amavis:[127.0.0.1]:10024
daemon_directory = /usr/libexec/postfix
debug_peer_level = 2
enable_server_options = yes
html_directory = no
inet_interfaces = all
mail_owner = postfix
mailbox_size_limit = 0
mailbox_transport = cyrus
mailq_path = /usr/bin/mailq
manpage_directory = /usr/share/man
mydestination = $myhostname,localhost.$mydomain,localhost,192.168.123.10
mydomain = LOCALDOMAIN.org
mydomain_fallback = localhost
myhostname = mail.LOCALDOMAIN.org
mynetworks = 127.0.0.0/8
mynetworks_style = host
newaliases_path = /usr/bin/newaliases
owner_request_special = no
queue_directory = /private/var/spool/postfix
readme_directory = /usr/share/doc/postfix
recipient_delimiter = +
sample_directory = /usr/share/doc/postfix/examples
sendmail_path = /usr/sbin/sendmail
setgid_group = postdrop
smtpd_pw_server_security_options = login
smtpd_recipient_restrictions = permit_sasl_authenticated,permit_mynetworks,reject_unauth_destination,permit
smtpd_sasl_auth_enable = yes
smtpd_tls_key_file =
smtpd_use_pw_server = yes
unknown_local_recipient_reject_code = 550
virtual_mailbox_domains = hash:/etc/postfix/virtual_domains
virtual_transport = lmtp:unix:/var/imap/socket/lmtp
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